Fellow 1986-1991

Area of Current Research Interests
Genetics of disease and variation, including pigmentation and eye disorders. Mouse models of human genetics.

Other Information
Genetics Society, Balfour Lecturer – 1986
Elected member of the Institute – 1996
President International Mammalian Genome Society – 2002-2004
Elected Fellow Royal Society of Edinburgh – 2008
Vice President, Genetics Society – 2008-2012
President, European Society for Pigment Cell Research – 2013-2016
Board Member, National Centre for the 3Rs – 2014-2019
Retired from MRC/University of Edinburgh – 2021
Professor Emeritus, University of Edinburgh – 2022